Numbers game.
Our Cyclone had an advertised tank capacity of 110 gallons. That worked out to two 50 gallon tanks and 10 gallons for the water heater.
I ended up replacing the water tanks for a 'capacity issue', 15-20 gallons of unusable water - and un-drainable as well.
The 50 gallon spec on the tanks calculated out to the outside measured dimensions of the tanks.
Take about 5 gallons away for interior dimensions (assuming 1/4" tank walls).
The vent line was on the sidewalls is near the top of the tanks. Once the water hits the vent you are done filling (by gravity means), so you can subtract another 1/2' of usable water. Take another 1/2" off the bottom and you lose 5-6 gallons of usable capacity.
Maybe the above can help explain your capacity issue.
The vent lines mentioned above were a frequent problem before the introduction of the can-leak valves.
My original vent lines drooped between the tanks and the vent in the docking stations. Once water got in the vent - it stopped venting.
Useless info for your issue.
We have a 30 gallon advertised fuel cell in the trailer (toy hauler). Again, outside measurements. Fuel tanks can't be filled 100%, need some air space. As a result, the most I ever got in the tanks was 25 gallons.
Doesn't seem like much, but is a 17% difference.
Cyclone water tanks
https://heartlandowners.org/showthread.php/11365-And-so-it-begins?p=247358&viewfull=1#post247358